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Interestingly, they proposed an additional link to oxidative stress according to superoxide dismutase upregulation that may be explained by mitophagy during nitrogen limitation (Venditti and Di Meo, 2020)

This restoration of tissue function could also be accompanied by a reduction in the biomarkers associated with cellular aging, suggesting a return toward a more balanced state of cell function

P., Boulting, G

Along with GSTs, some of these enzymes offer additional routes by which ascorbate and, particularly, glutathione can be linked to peroxide metabolism